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A marine habitat is a that supports . Marine life depends in some way on the that is in the sea (the term marine comes from the mare, meaning sea or ocean). A habitat is an or environmental area inhabited by one or more living .Abercrombie, M., Hickman, C.J. and Johnson, M.L. 1966. A Dictionary of Biology. Penguin Reference Books, London The marine environment supports many kinds of these habitats.

Marine habitats can be divided into and habitats. Coastal habitats are found in the area that extends from as far as the tide comes in on the out to the edge of the continental shelf. Most marine life is found in coastal habitats, even though the shelf area occupies only seven percent of the total ocean area. Open ocean habitats are found in the deep ocean beyond the edge of the continental shelf.

Alternatively, marine habitats can be divided into and . Pelagic habitats are found near the surface or in the open , away from the bottom of the ocean. habitats are near or on the bottom of the ocean. An organism living in a pelagic habitat is said to be a pelagic organism, as in . Similarly, an organism living in a demersal habitat is said to be a demersal organism, as in . Pelagic habitats are intrinsically shifting and ephemeral, depending on what are doing.

Marine habitats can be modified by their inhabitants. Some marine organisms, like , , and , are ecosystem engineers which reshape the marine environment to the point where they create further habitat for other organisms. By volume the ocean provides most of the habitable space on the planet. Living Ocean NASA Science. Retrieved 17 December 2016.


Overview
In contrast to terrestrial habitats, marine habitats are shifting and . Swimming organisms find areas by the edge of a continental shelf a good habitat, but only while bring nutrient rich water to the surface. Shellfish find habitat on sandy beaches, but storms, tides and currents mean their habitat continually reinvents itself.

The presence of is common to all marine habitats. Beyond that many other things determine whether a marine area makes a good habitat and the type of habitat it makes. For example:

  • temperature – is affected by geographical , , , the discharge of rivers, and by the presence of hydrothermal vents or
  • sunlight – processes depend on how deep and the water is
  • nutrients – are transported by ocean currents to different marine habitats from , or by upwellings from the deep sea, or they sink through the sea as
  • salinity – varies, particularly in or near , or by hydrothermal vents
  • dissolved gases – oxygen levels in particular, can be increased by wave actions and decreased during
  • acidity – this is partly to do with dissolved gases above, since the acidity of the ocean is largely controlled by how much carbon dioxide is in the water.
  • turbulence – , fast currents and the agitation of water affect the nature of habitats
  • cover – the availability of cover such as the adjacency of the , or the presence of floating objects
  • substrate – The slope, orientation, profile and of hard substrates, and particle size, sorting and density of sediment bottoms can make a big difference to the life forms that can settle on it.
  • the occupying organisms themselves – since organisms modify their habitats by the act of occupying them, and some, like corals, kelp, mangroves and seagrasses, create further habitats for other organisms.

[[File:BlueMarble-2001-2002.jpg|thumb|300px|right|Only 29 percent of the world surface is land. The rest is ocean, home to the marine habitats. The oceans are nearly four kilometres deep on average and are fringed with coastlines that run for nearly 380,000 kilometres.|alt=Two views of the ocean from space]]

There are five major oceans, of which the is nearly as large as the rest put together. Coastlines fringe the land for nearly 380,000 kilometres.

Pacific Ocean155.646.4679.649.64.3710.924135,663 CIA Factbook: Pacific ocean.
Atlantic Ocean76.822.9313.422.54.088.605111,866 CIA Factbook: Atlantic ocean.
Indian Ocean68.620.4269.319.63.937.25866,526 CIA Factbook: Indian ocean.
Southern Ocean20.36.191.56.74.517.23517,968 CIA Factbook: Southern ocean.
Arctic Ocean14.14.217.01.21.214.66545,389 CIA Factbook: Arctic ocean.
Overall335.3 1370.8Elert, Glenn Volume of Earth's Oceans. The Physics Factbook. Retrieved 19 April 2008. 4.0910.924377,412

Altogether, the ocean occupies 71 percent of the world surface, averaging nearly four kilometres in depth. By volume, the ocean contains more than 99 percent of the Earth's liquid water. Where is Earth's water?, United States Geological Survey.Eakins, B.W. and G.F. Sharman, Volumes of the World's Oceans from ETOPO1, National Geophysical Data Center, Boulder, CO, 2010. Water in Crisis: Chapter 2, Peter H. Gleick, Oxford University Press, 1993. The science fiction writer Arthur C. Clarke has pointed out it would be more appropriate to refer to the planet Earth as the planet Sea or the planet Ocean. Planet "Earth": We Should Have Called It "Sea" Quote Invertigator, 25 January 2017. Unveiling Planet Ocean NASA Science, 14 March 2002.

Marine habitats can be broadly divided into and habitats. habitats are the habitats of the open , away from the bottom of the ocean. Demersal habitats are the habitats that are near or on the bottom of the ocean. An organism living in a pelagic habitat is said to be a pelagic organism, as in . Similarly, an organism living in a demersal habitat is said to be a demersal organism, as in . Pelagic habitats are intrinsically ephemeral, depending on what are doing.

The land-based ecosystem depends on topsoil and fresh water, while the depends on dissolved nutrients washed down from the land.

Ocean deoxygenation poses a threat to marine habitats, due to the growth of low oxygen zones.


Ocean currents
In marine systems, have a key role determining which areas are effective as habitats, since ocean currents transport the basic nutrients needed to support marine life. Ocean Habitats Marietta College. Retrieved 17 April 2011. are the life forms that inhabit the ocean that are so small (less than 2 mm) that they cannot effectively propel themselves through the water, but must drift instead with the currents. If the current carries the right nutrients, and if it also flows at a suitably shallow depth where there is plenty of sunlight, then such a current itself can become a suitable habitat for photosynthesizing tiny algae called . These tiny plants are the primary producers in the ocean, at the start of the . In turn, as the population of drifting phytoplankton grows, the water becomes a suitable habitat for , which feed on the phytoplankton. While phytoplankton are tiny drifting plants, are tiny drifting animals, such as the of and marine invertebrates. If sufficient zooplankton establish themselves, the current becomes a candidate habitat for the that feed on them. And then if sufficient forage fish move to the area, it becomes a candidate habitat for larger and other marine animals that feed on the forage fish. In this dynamic way, the current itself can, over time, become a moving habitat for multiple types of marine life.

Ocean currents can be generated by differences in the density of the water. How dense water is depends on how saline or warm it is. If water contains differences in salt content or temperature, then the different densities will initiate a current. Water that is saltier or cooler will be denser, and will sink in relation to the surrounding water. Conversely, warmer and less salty water will float to the surface. Atmospheric winds and pressure differences also produces surface currents, and . Ocean currents are also generated by the gravitational pull of the sun and moon (), and seismic activity ().

The rotation of the Earth affects the direction ocean currents take, and explains which way the large circular rotate in the image above left. Suppose a current at the equator is heading north. The Earth rotates eastward, so the water possesses that rotational momentum. But the further the water moves north, the slower the earth moves eastward. If the current could get to the North Pole, the earth would not be moving eastward at all. To conserve its rotational momentum, the further the current travels north the faster it must move eastward. So the effect is that the current curves to the right. This is the . It is weakest at the equator and strongest at the poles. The effect is opposite south of the equator, where currents curve left.


Topography

Biomass
One measure of the relative importance of different marine habitats is the rate at which they produce biomass.
and 2,500
(2025). 9780716728290, Macmillan. .
Includes freshwater
2,0000.28Spalding, Mark, Corinna Ravilious, and Edmund Green. 2001. World Atlas of Coral Reefs. Berkeley, CA: University of California Press and UNEP/WCMC.0.56
2,000
1,800
125
(2025). 9780415217705, Routledge. .
311 39


Coastal
are dynamic environments which constantly change, like the ocean which partially shape them. The Earth's natural processes, including and sea level change, result in the , accretion and resculpturing of coasts as well as the flooding and creation of continental shelves and drowned river valleys.

The main agents responsible for deposition and along coastlines are , and . The formation of coasts also depends on the they are made of – the harder the rocks the less likely they are to erode, so variations in rock hardness result in coastlines with different shapes.

often determine the range over which is deposited or eroded. Areas with high tidal ranges allow waves to reach farther up the shore, and areas with lower tidal ranges produce deposition at a smaller elevation interval. The tidal range is influenced by the size and shape of the coastline. Tides do not typically cause erosion by themselves; however, can erode as the waves surge up river from the ocean.Davidson (2002), p.421.

Waves erode coastline as they break on shore releasing their energy; the larger the wave the more energy it releases and the more sediment it moves. Sediment deposited by waves comes from eroded cliff faces and is moved along the coastline by the waves. Sediment deposited by rivers is the dominant influence on the amount of sediment located on a coastline.Easterbrook (1999).

The classified coasts as primary or secondary.Shepard FP (1937) Revised "Classification of Marine Shorelines" The Journal of Geology, 45(6): 602–624.

  • Primary coasts are shaped by non-marine processes, by changes in the land form. If a coast is in much the same condition as it was when sea level was stabilised after the last ice age, it is called a primary coast. "Primary coasts are created by erosion (the wearing away of soil or rock), deposition (the buildup of sediment or sand) or tectonic activity (changes in the structure of the rock and soil because of earthquakes). Many of these coastlines were formed as the sea level rose during the last 18,000 years, submerging river and glacial valleys to form bays and fjords." Habitats: Beaches - Coasts Office of Naval Research. Retrieved 17 April 2011. An example of a primary coast is a , which forms when a river deposits soil and other material as it enters the sea.
  • Secondary coasts are produced by marine processes, such as the action of the sea or by creatures that live in it. Secondary coastlines include , , , , and .

Continental coastlines usually have a continental shelf, a shelf of relatively shallow water, less than 200 metres deep, which extends 68 km on average beyond the coast. Worldwide, continental shelves occupy a total area of about 24 million km2 (9 million sq mi), 8% of the ocean's total area and nearly 5% of the world's total area. Continental shelf areas Earth trends. Retrieved 25 February 2010. World The World Factbook, CIA. Retrieved 26 February 2010. Since the continental shelf is usually less than 200 metres deep, it follows that coastal habitats are generally , situated in the sunlit . This means the conditions for processes so important for primary production, are available to coastal marine habitats. Because land is nearby, there are large discharges of nutrient rich into coastal waters. Further, periodic from the deep ocean can provide cool and nutrient rich currents along the edge of the continental shelf.

As a result, coastal marine life is the most abundant in the world. It is found in , and , near sandy shores and rocky coastlines, around and on or above the continental shelf. include small forage fish as well as the larger that feed on them. thrive in inshore waters where high productivity results from and shoreline run off of nutrients. Some are partial residents that spawn in streams, estuaries and bays, but most complete their life cycle in the zone.Moyle and Cech, 2004, page 572 There can also be a mutualism between species that occupy adjacent marine habitats. For example, just below level have a mutually beneficial relationship with forests at high tide level and meadows in between: the reefs protect the mangroves and seagrass from strong currents and waves that would damage them or the sediments in which they are rooted, while the mangroves and seagrass protect the coral from large influxes of , fresh water and . This additional level of variety in the environment is beneficial to many types of coral reef animals, which for example may feed in the sea grass and use the reefs for protection or breeding.

(1989). 9780080377186, Routledge.

Coastal habitats are the most visible marine habitats, but they are not the only important marine habitats. Coastlines run for 380,000 kilometres, and the total volume of the ocean is 1,370 million cu km. This means that for each metre of coast, there is 3.6 cu km of ocean space available somewhere for marine habitats.


Intertidal
, those areas close to shore, are constantly being exposed and covered by the ocean's . A huge array of life lives within this zone.

Shore habitats range from the upper intertidal zones to the area where land vegetation takes prominence. It can be underwater anywhere from daily to very infrequently. Many species here are scavengers, living off of sea life that is washed up on the shore. Many land animals also make much use of the shore and intertidal habitats. A subgroup of organisms in this habitat bores and grinds exposed rock through the process of .


Sandy shores
Sandy shores, also called , are coastal shorelines where accumulates. Waves and currents shift the sand, continually building and eroding the shoreline. Longshore currents flow parallel to the beaches, making waves break obliquely on the sand. These currents transport large amounts of sand along coasts, forming spits, and . Longshore currents also commonly create , which give beaches some stability by reducing erosion. Habitats: Beaches - Characteristics Office of Naval Research. Retrieved 17 April 2011.

Sandy shores are full of life. The grains of sand host , and other microscopic creatures. Some fish and turtles return to certain beaches and spawn eggs in the sand. Birds habitat beaches, like , , , and . , such sea lions, recuperate on them. , , , , and are found on most beaches. Habitats: Beaches - Animal & Plant Life Office of Naval Research. Retrieved 17 April 2011.

Sand is a made from small grains or particles with diameters between about 60 μm and 2 mm.Wentworth CK (1922) "A scale of grade and class terms for clastic sediments" J. Geology, 30: 377–392. Mud (see mudflats below) is a sediment made from particles finer than sand. This small particle size means that mud particles tend to stick together, whereas sand particles do not. Mud is not easily shifted by waves and currents, and when it dries out, cakes into a solid. By contrast, sand is easily shifted by waves and currents, and when sand dries out it can be blown in the wind, accumulating into shifting . Beyond the high tide mark, if the beach is low-lying, the wind can form rolling hills of sand dunes. Small dunes shift and reshape under the influence of the wind while larger dunes stabilise the sand with vegetation.

Ocean processes grade loose sediments to particle sizes other than sand, such as or cobbles. Waves breaking on a beach can leave a , which is a raised ridge of coarser pebbles or sand, at the high tide mark. are made of particles larger than sand, such as cobbles, or small stones. These beaches make poor habitats. Little life survives because the stones are churned and pounded together by waves and currents.


Rocky shores
The relative solidity of seems to give them a permanence compared to the shifting nature of sandy shores. This apparent stability is not real over even quite short geological time scales, but it is real enough over the short life of an organism. In contrast to sandy shores, plants and animals can anchor themselves to the rocks. Tour of Rocky Shoreline Habitats Marietta College. Retrieved 17 April 2011.

Competition can develop for the rocky spaces. For example, can compete successfully on open intertidal rock faces to the point where the rock surface is covered with them. Barnacles resist desiccation and grip well to exposed rock faces. However, in the crevices of the same rocks, the inhabitants are different. Here can be the successful species, secured to the rock with their .

Rocky and sandy coasts are vulnerable because humans find them attractive and want to live near them. An increasing proportion of the humans live by the coast, putting pressure on coastal habitats.


Mudflats
are coastal wetlands that form when mud is deposited by tides or rivers. They are found in sheltered areas such as , , , and . Mudflats may be viewed as exposed layers of , resulting from deposition of , and marine animal . Most of the sediment within a mudflat is within the , and thus the flat is submerged and exposed approximately twice daily.


Mangrove forests and salt marshes
and form important coastal habitats in tropical and temperate areas respectively.

Mangroves are species of shrubs and medium size trees that grow in coastal sediment habitats in the and – mainly between ° N and ° S. The saline conditions tolerated by various species range from , through pure (30 to 40 ), to water concentrated by to over twice the salinity of ocean seawater (up to 90 ppt). There are many mangrove species, not all closely related. The term "mangrove" is used generally to cover all of these species, and it can be used narrowly to cover just mangrove trees of the genus .

Mangroves form a distinct characteristic saline or habitat, called a mangrove swamp or mangrove forest.Hogarth, Peter J. (1999) The Biology of Mangroves Oxford University Press, Oxford, England, What page? Mangrove swamps are found in depositional coastal environments, where fine sediments (often with high organic content) collect in areas protected from high-energy wave action. Mangroves dominate three quarters of tropical coastlines.


Estuaries
An is a partly enclosed body of with one or more or flowing into it, and with a free connection to the open .Pritchard, D. W. (1967) What is an estuary: physical viewpoint. p. 3–5 in: G. H. Lauf (ed.) Estuaries, A.A.A.S. Publ. No. 83, Washington, D.C. Estuaries form a transition zone between river environments and ocean environments and are subject to both marine influences, such as tides, waves, and the influx of saline water; and riverine influences, such as flows of fresh water and sediment. The inflow of both seawater and freshwater provide high levels of nutrients in both the water column and sediment, making estuaries among the most productive natural habitats in the world.McLusky, D.S. and Elliott, M. (2004) "The Estuarine Ecosystem: ecology, threats and management." New York: Oxford University Press Inc.

Most estuaries were formed by the flooding of river-eroded or glacially scoured valleys when sea level began to rise about 10,000-12,000 years ago.Wolanski, E. (2007) "Estuarine Ecohydrology." Amsterdam, the Netherlands: Elsevier. They are amongst the most heavily populated areas throughout the world, with about 60% of the world's population living along estuaries and the coast. As a result, estuaries are suffering degradation by many factors, including sedimentation from soil erosion from deforestation; overgrazing and other poor farming practices; overfishing; drainage and filling of wetlands; eutrophication due to excessive nutrients from sewage and animal wastes; pollutants including heavy metals, PCBs, radionuclides and hydrocarbons from sewage inputs; and diking or damming for flood control or water diversion.

Estuaries provide habitats for a large number of organisms and support very high productivity. Estuaries provide habitats for and nurseries,Bronwyn M. Gillanders, Evidence of connectivity between juvenile and adult habitats for mobile marine fauna: an important component of nurseries. 2003. Marine Ecology Progress Series as well as bird populations.Jennifer A. Gill, The buffer effect and large-scale population regulation in migratory birds. 2001. Nature 412, 436-438 Two of the main characteristics of estuarine life are the variability in and . Many species of and have various methods to control or conform to the shifts in salt concentrations and are termed and . Many animals also to avoid and to live in the more stable sedimental environment. However, large numbers of bacteria are found within the sediment which have a very high oxygen demand. This reduces the levels of oxygen within the sediment often resulting in partially conditions, which can be further exacerbated by limited water flux. are key primary producers in estuaries. They move with the water bodies and can be flushed in and out with the . Their productivity is largely dependent on the of the water. The main phytoplankton present are and which are abundant in the sediment.


Kelp forests
are underwater areas with a high density of . They form some of the most productive and dynamic on Earth.Mann, K.H. 1973. Seaweeds: their productivity and strategy for growth. Science 182: 975-981. Smaller areas of anchored kelp are called kelp beds. Kelp forests occur worldwide throughout and coastal oceans.

Kelp forests provide a unique three-dimensional habitat for marine organisms and are a source for understanding many ecological processes. Over the last century, they have been the focus of extensive research, particularly in ecology, and continue to provoke important ideas that are relevant beyond this unique ecosystem. For example, kelp forests can influence coastal patternsJackson, G.A. and C.D. Winant. 1983. Effect of a kelp forest on coastal currents. Continental Shelf Report 2: 75-80. and provide many ecosystem services.Steneck, R.S., M.H. Graham, B.J. Bourque, D. Corbett, J.M. Erlandson, J.A. Estes and M.J. Tegner. 2002. Kelp forest ecosystems: biodiversity, stability, resilience and future. Environmental Conservation 29: 436-459.

However, humans have contributed to kelp forest degradation. Of particular concern are the effects of nearshore ecosystems, which can release from their normal population regulation and result in the over-grazing of kelp and other algae.Sala, E., C.F. Bourdouresque and M. Harmelin-Vivien. 1998. Fishing, trophic cascades, and the structure of algal assemblages: evaluation of an old but untested paradigm. Oikos 82: 425-439. This can rapidly result in transitions to barren landscapes where relatively few species persist.Dayton, P.K. 1985a. Ecology of kelp communities. Annual Review of Ecology and Systematics 16: 215-245.

Frequently considered an ecosystem engineer, kelp provides a physical substrate and habitat for kelp forest communities.Jones, C.G., J. H. Lawton and M. Shachak. 1997. Positive and negative effects of organisms as physical ecosystem engineers. Ecology 78: 1946-1957. In algae (Kingdom: ), the body of an individual organism is known as a rather than as a plant (Kingdom: ). The morphological structure of a kelp thallus is defined by three basic structural units:

* The holdfast is a root-like mass that anchors the thallus to the sea floor, though unlike true roots it is not responsible for absorbing and delivering nutrients to the rest of the thallus;
* The stipe is analogous to a plant stalk, extending vertically from the holdfast and providing a support framework for other morphological features;
* The are leaf- or blade-like attachments extending from the stipe, sometimes along its full length, and are the sites of nutrient uptake and photosynthetic activity.
In addition, many kelp species have , or gas-filled bladders, usually located at the base of fronds near the stipe. These structures provide the necessary buoyancy for kelp to maintain an upright position in the water column.

The environmental factors necessary for kelp to survive include hard substrate (usually rock), high nutrients (e.g., nitrogen, phosphorus), and light (minimum annual dose > 50 E m−2Druehl, L.D. 1981. The distribution of Laminariales in the North Pacific with reference to environmental influences. Proceedings of the International Congress on Systematic Evolution and Biology 2: 248-256.). Especially productive kelp forests tend to be associated with areas of significant oceanographic , a process that delivers cool nutrient-rich water from depth to the ocean's . Water flow and turbulence facilitate nutrient assimilation across kelp fronds throughout the water column.Wheeler, W.N. 1980. Effect of boundary layer transport on the fixation of carbon by the giant kelp Macrocystis pyrifera. Marine Biology 56: 103-110. Water clarity affects the depth to which sufficient light can be transmitted. In ideal conditions, giant kelp ( Macrocystis spp.) can grow as much as 30-60 centimetres vertically per day. Some species such as Nereocystis are while others like Eisenia are , living for more than 20 years.Steneck, R.S. and M.N. Dethier. 1994. A functional group approach to the structure of algal-dominated communities. Oikos 69: 476-498. In perennial kelp forests, maximum growth rates occur during upwelling months (typically spring and summer) and die-backs correspond to reduced nutrient availability, shorter photoperiods and increased storm frequency.


Seagrass meadows
are from one of four plant families which grow in marine environments. They are called seagrasses because the leaves are long and narrow and are very often green, and because the plants often grow in large meadows which look like grassland. Since seagrasses and are submerged, they must grow submerged in the , where there is enough sunlight. For this reason, most occur in shallow and sheltered coastal waters anchored in sand or mud bottoms.

Seagrasses form extensive , which can be either monospecific (made up of one species) or multispecific (where more than one species co-exist). Seagrass beds make highly diverse and productive . They are home to such as juvenile and adult fish, and free-living and , , , and . Few species were originally considered to feed directly on seagrass (partly because of their low nutritional content), but scientific and improved working methods have shown that seagrass is a highly important link in the food chain, with hundreds of species feeding on seagrasses worldwide, including , , , , , , and .

Seagrasses are ecosystem engineers in the sense that they partly create their own habitat. The leaves slow down water-currents increasing , and the seagrass and stabilize the seabed. Their importance to associated species is mainly due to provision of shelter (through their three-dimensional structure in the water column), and due to their extraordinarily high rate of primary production. As a result, seagrasses provide coastal zones with ecosystem services, such as , protection, production and protection against coastal . Seagrass meadows account for 15% of the ocean's total carbon storage.


Reefs
A reef is a ridge or of rock, or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic processes but there are also reefs such as the of tropical waters formed by processes dominated by corals and . such as shipwrecks and other anthropogenic underwater structures may occur intentionally or as the result of an accident, and sometimes have a designed role in enhancing the physical complexity of featureless sand bottoms, thereby attracting a more diverse assemblage of organisms. Reefs are often quite near to the surface, but not all definitions require this. Fringing reefs, the most common type of reef, are found close to shorelines and surrounding islands.


Rocky reefs
Rocky reefs are underwater outcrops of rock projecting above the adjacent unconsolidated surface with varying relief. They can be found in depth ranges from to deep water and provide a substrate for a large range of sessile benthic organisms, and shelter for a large range of mobile organisms.


Coral reefs
comprise some of the densest and most diverse habitats in the world. The best-known types of reefs are which exist in most tropical waters; however, coral reefs can also exist in cold water. Reefs are built up by and other -depositing animals, usually on top of a rocky outcrop on the ocean floor. Reefs can also grow on other surfaces, which has made it possible to create . Coral reefs also support a huge community of life, including the corals themselves, their symbiotic , tropical fish and many other organisms.

Much attention in marine biology is focused on coral reefs and the El Niño weather phenomenon. In 1998, coral reefs experienced the most severe mass bleaching events on record, when vast expanses of reefs across the world died because sea surface temperatures rose well above normal.NOAA (1998) Record-breaking coral bleaching occurred in tropics this year. National Oceanic and Atmospheric Administration, Press release (October 23, 1998).ICRS (1998) Statement on Global Coral Bleaching in 1997-1998. International Coral Reef Society, October 15, 1998. Some reefs are recovering, but scientists say that between 50% and 70% of the world's coral reefs are now endangered and predict that could exacerbate this trend.Bryant, D., Burke, L., McManus, J., et al. (1998) "Reefs at risk: a map-based indicator of threats to the world's coral reefs". World Resources Institute, Washington, D.C.Goreau, T. J. (1992) "Bleaching and Reef Community Change in Jamaica: 1951 - 1991". Am. Zool. 32: 683-695.Sebens, K. P. (1994) "Biodiversity of Coral Reefs: What are We Losing and Why?" Am. Zool., 34: 115-133Wilkinson, C. R., and Buddemeier, R. W. (1994) "Global Climate Change and Coral Reefs:Implications for People and Reefs". Report of the UNEP-IOC-ASPEI-IUCN Global Task Team on the Implications of Climate Change on Coral Reefs. IUCN, Gland, Switzerland.


Surface waters

Surface microlayer
The surface microlayer of the ocean serves as the transitional area between the atmosphere and the ocean. It covers around 70% of the Earth's surface as it covers most of the ocean waters on the planet. The microlayer is known for its unique biological and chemical properties which give it a small ecosystem of its own and serves as a distinct habitat from the deeper ocean waters.

The surface microlayer is not in fact entirely aqueous like the rest of the ocean, but is closer to a kind of hydrated gel composed of concentrated nutrients forming a biological film over the water it covers. This film is rich in microbes which mediate the interactions between the sun, the atmosphere, and the waters below.

Although thin, the surface microlayer is critical for life beneath it. Because of the environment rich in microbes and nutrients, larvae of fish and other aquatic animals are often laid in the microlayer to incubate. The plankton in the microlayer are distinctly adapted to withstand high levels of radiation, and serve as buffers to prevent this potentially harmful radiation from reaching the deeper water. Environmental changes such as aerosols or dust storms can cause these surface plankton to become overproductive, leading to . Because of the unique properties of the microlayer, pollutants often accumulate within and use it to reach other parts of the ocean. Hydrophobic compounds, such as , flame retardants, and heavy metals, have a particular affinity for the surface microlayer. Recently, the abundance of aerosols and has also had an impact on the SML and their accumulation has led to many problems, such as animal ingestion of these compounds leading to widespread disruption of balance and spread of these compounds among marine communities.

The surface microlayer is also critical to gas exchange between the atmosphere and the ocean. Because the microlayer is filled with microbes, it is widely theorized that it plays a critical role in gas exchange and uptake of nutrients, but relatively little data on this has been collected. The central feature of the microlayer is the temperature, as it is an indicator of how pollutants and human activity affects the ocean.


Epipelagic zone
The surface waters are sunlit. The waters down to about 200 metres are said to be in the . Enough sunlight enters the epipelagic zone to allow by . The epipelagic zone is usually low in nutrients. This partially because the organic debris produced in the zone, such as excrement and dead animals, sink to the depths and are lost to the upper zone. Photosynthesis can happen only if both sunlight and nutrients are present. Ocean ecology: sunlit surface waters WWF. Retrieved 17 May 2011.

In some places, like at the edge of continental shelves, nutrients can from the ocean depth, or can be distributed by storms and ocean currents. In these areas, given that both sunlight and nutrients are now present, phytoplankton can rapidly establish itself, multiplying so fast that the water turns green from the chlorophyll, resulting in an . These nutrient rich surface waters are among the most biologically productive in the world, supporting billions of tonnes of biomass.

"Phytoplankton are eaten by - small animals which, like phytoplankton, drift in the ocean currents. The most abundant zooplankton species are and : tiny that are the most numerous animals on Earth. Other types of zooplankton include and the of fish, , , and other marine organisms". In turn, the zooplankton are eaten by animals, including some , small like herrings and sardines, , , and the largest animal in the world, the . Yet again, moving up the , the small forage fish are in turn eaten by larger predators, such as tuna, marlin, sharks, large squid, seabirds, dolphins, and .


Open ocean
The open ocean is relatively unproductive because of a lack of nutrients, yet because it is so vast, it has more overall primary production than any other marine habitat. Only about 10 percent of marine species live in the open ocean. But among them are the largest and fastest of all marine animals, as well as the animals that dive the deepest and migrate the longest. In the depths lurk animal that, to our eyes, appear hugely alien. Blue Planet: Open ocean WWF. Retrieved 17 May 2011.


Deep sea
The starts at the , the point where sunlight loses most of its energy in the water. Many life forms that live at these depths have the ability to create their own light a unique evolution known as .

In the deep ocean, the waters extend far below the epipelagic zone, and support very different types of pelagic life forms adapted to living in these deeper zones.Moyle and Cech, 2004, page 585

Much of the 's energy is supplied by the open ocean in the form of . In deep water, is a continuous shower of mostly organic detritus falling from the upper layers of the water column. Its origin lies in activities within the productive . Marine snow includes dead or dying , (), fecal matter, sand, soot and other inorganic dust. The "snowflakes" grow over time and may reach several centimetres in diameter, travelling for weeks before reaching the ocean floor. However, most organic components of marine snow are consumed by , and other filter-feeding animals within the first 1,000 metres of their journey, that is, within the epipelagic zone. In this way marine snow may be considered the foundation of deep-sea and : As sunlight cannot reach them, deep-sea organisms rely heavily on marine snow as an energy source.

Some deep-sea pelagic groups, such as the , , marine hatchetfish, and families are sometimes termed pseudoceanic because, rather than having an even distribution in open water, they occur in significantly higher abundances around structural oases, notably and over continental slopes. The phenomenon is explained by the likewise abundance of prey species which are also attracted to the structures.

The fish in the different pelagic and deep water benthic zones are physically structured, and behave in ways, that differ markedly from each other. Groups of coexisting species within each zone all seem to operate in similar ways, such as the small mesopelagic vertically migrating plankton-feeders, the bathypelagic , and the deep water benthic . "Moyle and Cech, 2004, p. 591

species, with spiny fins, are rare among deep sea fishes, which suggests that deep sea fish are ancient and so well adapted to their environment that invasions by more modern fishes have been unsuccessful.Haedrich RL (1996) "Deep-water fishes: evolution and adaptation in the earth's largest living spaces" Journal of Fish Biology 49(sA):40-53. The few ray fins that do exist are mainly in the and , which are also ancient forms. Most deep sea pelagic fishes belong to their own orders, suggesting a long evolution in deep sea environments. In contrast, deep water benthic species, are in orders that include many related shallow water fishes.Moyle and Cech, 2004, page 586

The is a deep sea eel with an enormous loosely hinged mouth. It can open its mouth wide enough to swallow a fish much larger than itself, and then expand its stomach to accommodate its catch.

(1998). 9780125476652, Academic Press.


Sea floor

Vents and seeps
Hydrothermal vents along the spreading centers act as , as do their opposites, . Such places support unique and many new marine microorganisms and other lifeforms have been discovered at these locations.


Trenches
The deepest recorded measure to date is the , near the , in the at 10,924 m (35,838 ft). At such depths, is extreme and there is no sunlight, but some life still exists. A white , a shrimp and a jellyfish were seen by the American crew of the Trieste when it dove to the bottom in 1960. Seven Miles Down: The Story of The Bathyscaph Trieste. , Rolex Deep Sea Special, January 2006.


Seamounts
Marine life also flourishes around that rise from the depths, where fish and other sea life congregate to spawn and feed.


Anthropogenic impacts
Mudflats are typically important regions for wildlife, supporting a large population, although levels of biodiversity are not particularly high. They are of particular importance to as well as crabs, shrimp, and shellfish. These areas along the coast act as a nursery for these animals by providing an area for reproduction and feeding. However, this can pose as an issue due to the high trafficking of the birds migrating for nesting, then leaving to return to their seasonal homes. Whatever pollutants the birds take in while breeding are brought back with them to their next location, thus polluting that area as well. In the mudflats have been classified as a Biodiversity Action Plan priority habitat. European countries such as France have also found it beneficial to use the Marine Influence Index (MII) to be able to monitor the responses to pollution the local plant and animal species may have as well as monitor any type of deviation from the natural patterns displayed previously.

Although many parts of the seafloor have yet to be explored, researchers have found that parts of it have been greatly affected by human activity. Bottom trawling, microplastic pollution, and industrial metals have slowly changed and altered the composition of the sea floor. refers to a commercial deep sea fishing technique in which the equipment drags across the sea floor. This has had an adverse effect on the seafloor as it changes the surface structure and composition. In addition, pollution has become an increasing problem to the seafloor as plastics and other debris are found in many of the sediments. Due to the build up of litter, the habitats and environments of organisms on the seafloor are being impacted and changed. This includes industrial facilities dumping new metals and minerals, such as , onto the seafloor that change the chemical composition of the water and poison the inhabitants.

File:Microplastic.jpg|Microplastics found in sediments on the seafloor File:Eurythenes plasticus (10.11646-zootaxa.4748.1.9) Figure 3.jpg|640 μm microplastic found in the deep sea amphipod Eurythenes plasticus

There are also negative anthropogenic impacts on deep sea habitats, including trash pollution and chemical pollution. Plastic pollution in particular, is one of the greatest forms of uncontrolled human activity that is visible in our oceans today. Researchers in the Northwestern south China Sea recorded large plastic-dominated litter piles in . These durable plastics can diffuse into smaller organisms and are then inadvertently consumed by humans in the food we eat and water we drink. Another threat to organisms lurking in the deep ocean is ghost fishing, and . Ghost fishing is the term that refers to any abandoned fishing gear in the ocean that continues to entangle and trap marine organisms. for example, have been recorded tangled around deep sea corals and continue ghost fishing for extended periods of time.

==Gallery==


See also
  • Effects of climate change on oceans
  • Future of Marine Animal Populations
  • Seashore wildlife
  • Underwater habitat (underwater habitats for humans)


Sources
  • Kritzer JP; Sale PF (2006). Marine metapopulations. Academic Press. .
  • Moyle, PB and Cech, JJ (2004) Fishes, An Introduction to Ichthyology. 5th Ed, Benjamin Cummings. .
  • Nybakken JW; Bertness MD (2005) Marine biology: an ecological approach. Sixth edition. Pearson/Benjamin Cummings. – organized by habitat, not classification


External links

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